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Can you hook up the fuel pressure gauge so it is "tee'd" into the supply line? That way you can observe if the fuel pressure remains adequate. I suspect the fuel rail is getting initial pressurization when you turn the key to 'start' but it is not remaining pressurized after the engine starts (and the key is not in the 'start' position). This would point to the drive relay not supplying power to the pumps.
Have you tested the two flywheel sensors?
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